BRITISH BRYOLOGICAL SOCIETY
Survey of the
Bryophytes of Arable Land
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Bryophytes associated with arable land
(from Fields of Vision Conference, Cambridge
2000, Ron Porley)
| Species | Status | Asexual | Sporophytes | Sexuality | Frequency of occurrence in arable |
| Liverworts and hornworts | . | . | . | . | . |
| Anthoceros agrestis | . | . | common | mono | often in arable but uncommon |
| Anthoceros punctatus | . | . | common | mono | often in arable but very local |
| Blasia pusilla | . | gemmae (two types) | occasional | dioi | occassionally recorded from arable |
| Fossombronia caespitiformis | NS | . | frequent | mono | most records from arable but rare |
| Fossombronia pusilla | . | . | frequent | mono | occassionally recorded from arable |
| Fossombronia wondraczeckii | . | . | common | mono | occassionally recorded from arable |
| Lophocolea bidentata | . | . | common | auto | opportunistic and not typically an arable species |
| Lunularia cruciata | . | gemmae | very rare | dioi | opportunistic and not typically an arable species |
| Phaeoceros carolinianus | EN | . | common | mono | occurs in arable but very rare |
| Phaeoceros laevis | . | perennating tubers | frequent | dioi | occasionally found in arable but uncommon |
| Riccia cavernosa | NS | . | common | mono | typical of damp arable but uncommon |
| Riccia crystallina | Intro | . | common | mono | characteristic of bulb & potato fields on Scilly |
| Riccia glauca | . | . | common | mono | a characteristic arable species |
| Riccia sorocarpa | . | . | common | mono | a characteristic arable species |
| Riccia subbifurca | NS | . | common | mono | uncommon in arable |
| Sphaerocarpos michelii | NS | . | frequent | dioi | characteristic plant of arable but uncommon |
| Sphaerocarpos texanus | VU | . | frequent | dioi | characteristic plant of arable but rare |
| Mosses | . | . | . | . | . |
| Acaulon muticum | . | . | abundant | auto or dioi | scarce and only occassional in arable |
| Aphanorhegma patens | . | . | abundant | paro or syno | rare in arable |
| Atrichum undulatum | . | . | common | mono | opportunistic and not typically an arable species |
| Barbula convoluta | . | tubers, Pcul | occasional | dioi | frequent in arable on calcareous soils |
| Barbula unguiculata | . | Pcul | occasional | dioi | frequent in arable on calcareous soils |
| Brachythecium rutabulum | . | . | common | auto | opportunistic and occurrence variable |
| Brachythecium velutinum | . | . | common | auto | opportunistic and occurrence variable |
| Bryum argenteum | . | bulbils, Pcul | occasional | dioi | common in arable |
| Bryum bicolor | . | bulbils, Pwild, tubers | frequent | dioi | a characteristic arable species |
| Bryum gemmiferum | . | bulbils, tubers (cul) | occasional | dioi | scarce in arable |
| Bryum gemmilucens | . | bulbils, Pcul | unknown | dioi | typically found in arable but very rare |
| Bryum klinggraeffii | . | tubers, Pwild+cul | unknown | dioi | a characteristic arable species |
| Bryum riparium | . | tubers | unknown | ? | in arable within flood zone of streams but rare |
| Bryum rubens | . | tubers, Pwild+cul | occasional | dioi | a characteristic arable species, often abundant |
| Bryum ruderale | . | tubers, Pwild+cul | unknown | dioi | not typically found in regularly disturbed arable |
| Bryum sauteri | . | tubers, Pcul | unknown | dioi | occasionally found in arable but uncommon |
| Bryum subapiculatum | . | tubers, Pcul | frequent | dioi | a characteristic arable species |
| Bryum violaceum | . | tubers | unknown | dioi | occasionally found in arable but uncommon |
| Ceratodon purpureus | . | Pwild+cul | frequent | dioi | common in arable |
| Chenia leptophylla | Intro | tubers | unknown | dioi | 3 British records, 1 in arable |
| Dicranella schreberiana | . | tubers | occasional | dioi | a characteristic arable species |
| Dicranella staphylina | . | tubers, Pcul | unknown | dioi | a characteristic arable species |
| Dicranella rufescens | . | tubers | occasional | dioi | uncommon in arable |
| Dicranella varia | . | tubers | frequent | dioi | common in arable |
| Didymodon fallax | . | Pcul | occasional | dioi | uncommon in arable |
| Didymodon tomaculosus | NT/BAP | tubers | unknown | dioi | most records from arable but very rare |
| Didymodon vinealis | . | Pcul | rare | dioi | rare in arable, perhaps overlooked |
| Ditrichum cylindricum | . | tubers | very rare | dioi | a characteristic arable species |
| Ditrichum pusillum | NS | tubers, Pcul | unknown* | dioi | rare in arable |
| Entosthodon fasicularis | . | . | frequent | auto | a characteristic arable species but uncommon |
| Ephemerum recurvifolium | NS | . | frequent | pseudodioi | uncommon in arable |
| Ephemerum serratum var. minutissimum | . | . | frequent | pseudodioi | occurrence sporadic but typically in arable |
| Ephemerum stellatum | EN/BAP | . | frequent | pseudodioi | 5 British records, 2 recently in arable |
| Eurhynchium hians | . | Pcul | rare | dioi | opportunistic and occurrence variable |
| Eurhynchium praelongum | . | . | frequent | dioi | opportunistic and occurrence variable |
| Fissidens bryoides | . | . | abundant | auto (rarely syno) | occasional in arable |
| Fissidens incurvus | . | . | abundant | dioi or auto | occasional in arable |
| Fissidens taxifolius | . | tubers | frequent | auto | occasional in arable |
| Fissidens viridulus | . | . | frequent | dioi, auto or syno | rare in arable |
| Funaria hygrometrica | . | Pwild+cul | frequent | auto | common in arable |
| Hennediella stanfordensis | NS | tubers | very rare | dioi | rare in arable |
| Leptobryum pyriforme | . | tubers | abundant | syno | occasional in arable |
| Microbryum curvicolle | . | . | abundant | auto | occasional in arable |
| Microbryum floerkeanum | NS | . | abundant | auto | often in arable but local |
| Microbryum davallianum | . | . | abundant | auto | occasional in arable |
| Microbryum starckeanum | . | . | abundant | auto | occasional in arable |
| Microbryum rectum | . | . | abundant | auto | occasional to frequent in arable |
| Physcomitrium pyriforme | . | . | abundant | auto | occasional in arable |
| Plagiomnium rostratum | . | . | frequent | syno | opportunistic and occurrence variable |
| Pleuridium acuminatum | . | . | abundant | paro | occasional in arable |
| Pleuridium subulatum | . | . | abundant | auto | occasional in arable |
| Pohlia camptotrachela | . | bulbils | unknown | dioi | rare in arable |
| Pohlia melanodon | . | tubers | occasional | dioi | uncommon in arable |
| Pohlia lescuriana | . | tubers | rare | dioi | uncommon in arable |
| Pohlia lutescens | . | tubers | unknown | dioi | uncommon in arable |
| Pterygoneurum ovatum | NS | . | frequent | auto | rare in arable |
| Pseudephemerum nitidum | . | . | abundant | syno | uncommon in arable |
| Pseudocrossidium hornschuchianum | . | Pcul | rare | dioi | rare in arable |
| Rhytidiadelphus squarrosus | . | . | rare | dioi | opportunistic and occurrence variable |
| Tortula acaulon | . | Pcul | abundant | auto | a characteristic arable species |
| Tortula acaulon var. papillosa | . | . | abundant | auto | a characteristic arable species but rare |
| Tortula acaulon var. schreberiana | . | . | abundant | auto | a characteristic arable species but rare |
| Tortula modica | . | . | abundant | auto | uncommon in arable |
| Tortula truncata | . | . | frequent | auto | a characteristic arable species |
| Weissia brachycarpa | . | . | abundant | auto | occasional in arable |
| Weissia longifolia var.angustifolia | . | . | abundant | auto | rare in arable |
| Weissia longifolia var. longifolia | . | . | abundant | auto | uncommon in arable |
| Weissia mittenii | EX | . | common | auto | 5 British localities, not all in arable |
| Weissia multicapsularis | VU/BAP | . | abundant | auto | very rare, recent records on Cornish hedges within arable |
| Weissia rostellata | NT | . | abundant | auto | rare in arable |
| Weissia rutilans | . | . | abundant | auto | rare in arable |
| Weissia squarrosa | EN | . | abundant | auto | rare in arable |